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An interesting thing I found in Palm Tree Paradise...
Diamond City Mayor
Diamond City Leader
I disabled layer 0 in this Wario Land 4 level, and look what I found:
Yes, those are the debug level tiles underneath the grass ones. Was this level in development before the graphics were finalised?
Huh, must've been! Did you find anything of the sort in levels like Hall of Hieroglyphs? Maybe this could shed some light on what order the levels were made in if it appears in more.
Diamond City Mayor
Diamond City Leader
Nothing in the Hall of Hieroglyphs. Just in this one room in Palm Tree Paradise.